Forest Regeneration
Often it is desirable to cut or regenerate
overstory trees in hardwood forests in order to encourage the young, thick
vegetation required as cover and food for wildlife such as ruffed grouse,
rabbits, deer, blue-winged warblers, rufous-sided towhees, and white-eyed
vireos. For instance, ruffed grouse require areas of young forest consisting of
between 2000 to 8000 saplings at least 5 feet tall per acre. This dense,
vertically oriented vegetation provides needed cover that is lacking in older,
more open woods. Unfortunately, many people have a negative opinion of cutting
trees. While timber cutting can have adverse environmental impacts when done in
an irresponsible manner, the responsible harvesting of trees can be an important
method of managing forestland for wildlife as well as for forest products. This
Habitat How-To is primarily aimed at managing forestland for wildlife habitat.
Landowners whose main objective is managing their forests for timber production
should seek technical guidance from the Kentucky Division of Forestry
(1-800-866-0555).
Another equally important reason to regenerate a
hardwood stand is to encourage the regrowth of desirable tree species such as
oaks and other hard mast producers. While all of our native tree species have
important ecological functions, oaks are the dominant component of most of
Kentucky’s hardwood forests. Oak trees also provide acorns, which are one of
the most important food sources for many of our wildlife species. However, in
many areas forests have been degraded by high-grading, a timber harvesting
method which removes all of the productive trees and leaves only poor quality
trees of less desirable species. The low quality trees remaining after such
harvests are detrimental because they shade the ground and prevent oaks,
hickories, and other hard mast producing trees from growing back. When the large
overstory trees in a forest stand are cut correctly, sunlight can reach the
forest floor and stimulate the regrowth, or "regeneration" of a new,
healthier forest.
Sprouts and Seeds: The Birth of a
New Forest
Young trees that grow up to regenerate a forest
after the overstory trees have been removed can come from three different
sources: 1) stump sprouting, 2) advance reproduction, and 3) seeds. It is
important to recognize which source is likely to be the most successful on a
particular site in order to determine the most appropriate method of
regeneration. Much of this discussion focuses on regenerating oak species. This
is because, as previously mentioned, oaks provide an important food source for
many wildlife species, but also because oaks are shade intolerant species. Shade
intolerant species, like oaks, need lots of sunlight to regenerate, unlike shade
tolerant species such as maples, which grow well under a closed forest canopy.
This means that for oaks and other shade tolerant species to persist in a
forest, there must be periodic natural disturbances or intentional human
management to open up the canopy and let in sunlight. Shade tolerant species
will be present regardless of whether or not the forest is managed.
Stump sprouting usually accounts for most of the
saplings which grow back as a result of clear-cutting stands where the majority
of the overstory trees are less than 16 inches in diameter at breast height (d.b.h.).
Stump sprouts are desirable when attempting to regrow trees of the same species
as those being cut. Stump sprouts will grow much faster than seedlings because
they already have an established root system. To encourage stump sprouting, all
overstory trees should be cut between November and March leaving stumps less
than 12 inches above the ground.
Saplings already present in the understory prior
to a timber harvest account for the second source of new tree growth. These
saplings are collectively called advance reproduction. Advance reproduction of
oaks or other desirable species is essential for successfully regenerating
hardwood stands where stump sprouting is not likely. A minimum of 400 oak
seedlings at least 4.5 feet tall per acre is necessary to successfully
regenerate an oak-dominated forest stand. Suitable densities of oak advance
reproduction often occur one to two years following a good year of acorn
production. If adequate numbers of saplings are not available one year due to
poor acorn crops during previous years, and there are healthy, mature oak trees
present, the harvest should be delayed until the density of saplings is
improved.
Advance reproduction of oaks tends to be most
successful on drier south and west facing slopes where there is less competition
from other tree species such as maples and poplar. When adequate advance
reproduction is present, the stand should be regenerated by clear-cutting.
Clear-cutting involves removing all trees in the stand larger than 1 inch d.b.h.,
except for advance reproduction of oaks or other desirable species. This timber
harvest technique provides high levels of sunlight to help the young oak sprouts
compete with other less desirable trees that grow best in the shade. A
modification of this method, which will benefit wildlife, is to leave some
clumps of soft mast producing shrubs such as dogwoods, serviceberry, viburnums,
and a few grapevines.
If advance oak reproduction is not present due to
a low number of mature oak trees, the stand should be regenerated with a 2 to 3
stage shelterwood cut. This method of harvesting removes approximately 30 –
40% of the canopy including suppressed or inferior oaks and all overstory trees
other than mature, healthy oaks. The goal of this step is to encourage heavy
acorn production from these healthy oak trees that are left, and at the same
time promote favorable conditions for those seeds to sprout and grow rapidly by
reducing competition from other trees. Once adequate advance reproduction has
resulted from the first partial cut, the remaining large overstory trees can
then be harvested. A shelterwood cut may take as long as 20 – 30 years to
complete but the initial thinning will produce a thicker understory desirable as
wildlife cover. On some highly productive sites, additional Timber Stand
Improvement (TSI) practices may be required along with the shelterwood cut to
successfully produce adequate oak regeneration. On some sites, oak regeneration
may not even be practical.
Distribution and Size of
Regeneration Cuts
The overall goal of hardwood forest
regeneration should be to maintain forestland in various aged stands to provide
for the different habitat requirements of a variety of wildlife. It is most
desirable for an oak forest to be composed of trees from both the red oak and
white oak groups. This is to assure both a good supply of acorns in years when
one type might have a complete failure to produce seeds, and a relatively
consistent food source through fall and winter (white oak acorns tend to drop
earlier). At least 40% or more of the total forestland should be old enough to
produce a reliable source of hard mast. Oak trees may begin producing acorns
between 20 to 30 years of age but do not reach optimum acorn production until
they are about 50 years old. In order to maintain at least 40% of the total
forested area in trees 30 years old or older, no more than 10% of the forest
should be regenerated within a 5 year period, assuming that you are starting
with mature forest. This may be unrealistic if you are trying to remedy the
effects of past high-grading and therefore have few mast producing trees at the
start of the operation.
Regeneration cuts of 5 – 20 acres are most
desirable, but even cuts of one acre or less can be beneficial on smaller
properties. These cuts should be scattered throughout the forested area with at
least 600 feet between cuts of the same age and should be interspersed with
areas of mature forest. Another option would be to concentrate regeneration cuts
around the edges of woodlands and leave a core area of mature trees within the
forest. This type of management would ensure a gradual transition (feathered
edge*) between forestland and surrounding habitats and reduce the negative
effects of abrupt edges such as increased nest predation.
Additional Management
Considerations
Tree tops and other slash produced during a
regeneration cut should be gathered into brush piles* which will provide cover
for small wildlife. When left scattered throughout the forest at ground level,
this slash can restrict the vision and movement of wildlife species such as
ruffed grouse, making them more vulnerable to predators. Log landings and skid
trails produced during forest management should be seeded to a legume* such as
white clover or Korean lespedeza and on steep areas orchard grass or other
desirable cool season grasses* (other than fescue) should be added to the mix to
reduce erosion. These legumes will not only provide additional forage for
wildlife but will also attract insects needed by young grouse, turkeys, and
other birds.
In mature hardwood stands, forest regeneration
can be accomplished as a by-product of harvesting marketable timber.
Regenerating hardwood stands that have been high-graded will probably not
produce marketable timber, but it may be a good source of firewood. In either
case, Best Management Practices (BMPs) should be used to prevent erosion and
other potential environmental concerns. In the years following a regeneration
cut, the forest may require additional management or Timber Stand Improvement (TSI)*.
The Kentucky Division of Forestry can provide you with information about
appropriate BMP’s for your operation. They can also provide additional
technical assistance in planning and implementing regeneration cuts, as well as
information on other forest management options.
*Related Habitat How-To references: